This hold (city isn't really the right description) exists within the forest known as The Felyal on the south coast of the Lowlands. It is spread organically throughout the trees without disturbing the natural environment, and at times it can be very hard to identify where the forest ends and the buildings begin.
Felyal is between the Ant city of Tark and the beetles in Collegium. This put it straight in the path of the Wasp Imperial Fourth Army. The Mantis proved the prowess of their weaponmasters during the destruction of that army, but were unable to stand against a renewed assault that knew what to expect.